4.88's and 33's a good combo?

[Cackalak Han]

Unless you have numbers from a similar vehicle, I'm not sure I would take that bet. Sure, you will turning higher rpm with bigger gears, but the engine will not be working as hard. Just because you need to keep it under 2500 rpm for 18 mpg with your 4.56 gears does not mean that will hold true with 5.29. Check in with Darren or ask your question on a forum with more modded 4Runners. All we can do here is speculate.

I heard from this guy who swapped the stock gear for 4.9FD on his Honda. Cruising at 80MPH went from 4k to 4.5k rpm. However, he noticed an INCREASED mpg. His speculation was that the engine didn't have to work as hard to cut through the air with the 4.9's. Same might hold true here.